52.Different paths, similar pressures: divergent drivers of genetic diversity despite convergent genomic signatures of selection in response to urban intensity in two oligolectic bee species

Molecular Ecology (2026), no.9: e70370

Baltz L., Gardein H., Greil H., Paxton R., Theodorou, P. Full text

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51. Semi-natural habitat cover, edge density and landscape conservation context shape the occupancy, species richness and reproductive performance of cavity-nesting Hymenoptera in an agricultural landscape

Landscape Ecology (2026) 41, 61

Wild C., Zelychenko J., Hopfenmüller S., Paxton R. Theodorou P. Full text

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50. Advancing pollinator science: A new global and integrative research platform

Advances in Pollinator Research (2026): 1: e188033

Beaurepaire A, Carneiro LT, Chakrabarti P, Disayathanoowat T, Erler S, Geppert C, Gonçalves LT, Kohl PL, Kurze C, Librán-Embid F, Osterman J, Parkinson RH, Prado A, Rutschmann B, Theodorou P, Villamar-Bouza L, Metodiev T, Stoev P, Warrit N, Straub L. Full text

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2025 (7)

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49. Genetic mapping in the red mason bee implicates ANTSR as an ancient sex-determining locus in bees and ants

PLOS Biology (2025), 23(11): e3003458

Rönneburg T., Taliadoros D., Anna Olsson A., Magnusson S., Huser L., Muhammad Fuad M., Everitt T., C. Martín-Hernández G., Theodorou P., Cederberg B., Paxton R., Seidelmann K., Webster M. Full text

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48. Integrating population genomics and historical occurrence data to assess the impact of land-use changes on Centris oil bees in the Brazilian Cerrado

Molecular Ecology (2025), e70153

Paulo S., Theodorou P., Martins A., Aquiar A., Shibata A., Kahnt B. Full text

 

47. Varroa destructor weakens the external immunity of honey bees (Apis mellifera) by impairing melittin production

Scientific Reports (2025), 15, 30623

Pusceddu M., Tragust S., Theodorou P., Ciabattini I., Navarro J., Corrias F., Atzei A., Angioni A., Floris I., Satta A. Full text

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46. Local floral resources and edge density within the urban ecosystem promote larger and less variable body size in the great banded furrow bee, Halictus scabiosae

BMC Ecology and Evolution (2025), 25, 75

Baltz L., de Vastey J., Gardein H., Klaus F., Greil H., Paxton R., Theodorou, P. Full text

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45. Metabarcoding assessment of arthropod diversity on green roofs in the metropolitan city of Hamburg

Journal of Urban Ecology (2025), 11, juaf003

Nevermann S., Hoffmann H., Theodorou P., Peters I., Husemann M. Full text

 

44. Prey and prejudice: Predation by the European bee-eater (Merops apiaster) has species-specific effects on the ecology and genetics of bumblebees

Oikos (2025), e11177

Kahnt B., Etel A., Jarosch F., Schweiger O., Çelikgil A., Theodorou P. Full text

 

43. Protected areas do not outperform urban wastelands in supporting insect pollinators and pollination in central Germany

Basic and Applied Ecology (2025), 84, 29-39

Theodorou P., Osterman W.H.A, Mrozek J.H., Wild B.S., Beckmann M., Osterman J., Paxton R. Full text

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​2024 (2)

 

42. Organic agriculture and annual flower strips reduce parasite prevalence in honey bees and boost colony growth in agricultural landscapes

Journal of Applied Ecology (2024), 61, 2146–2156 

Pluta P., Czechofsky K., Hass A., Frank L., Westerhoff A., Klingenberg H., Theodorou P., Westphal C., Paxton R. Full text 

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41. How seasonality, semi-natural habitat cover and compositional landscape heterogeneity affect pollen collection and development of Apis mellifera colonies in Mediterranean agro-sylvo-pastoral systems

Landscape Ecology (2024), 39, 26

Satta A., Lezzeri M., Brundu G., Floris I., Palmieri N., Pantaleoni A., Theodorou P.*, Pusceddu M. Full text 

*Corresponding author

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2023 (6)

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40. Global taxonomic, functional, and phylogenetic diversity of bees in apple orchards 

Science of the Total Environment  (2023), 901

Leclercq, N,........Theodorou P.,.......Vereecken, N.J. Full text

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39. Mason bees and honey bees synergistically enhance fruit set in sweet cherry orchards

Ecology and Evolution (2023),13, e10289

Osterman J., Benton F., Hellström S., Luderer-Pflimpfl M., Pöpel-Eisenbrandt A., Wild B., Theodorou P., Ulbricht C., Paxton R. Full text

 

38. Trouble in the tropics: pathogen spillover is a threat for native stingless bees

Biological Conservation (2023), 284, 110150

Fleites Ayil F. A., Medina M., Luis A., Quezada-Euán J., Stolle E., Theodorou P., Tragust S., Paxton R. Full text 

 

37. When lizards shift to a more plant-based lifestyle: The macroevolution of mutualistic lizard-plant-interactions (Squamata: Sauria/Lacertilia)

Molecular Phylogenetics and Evolution (2023), 186

Kahnt B., Theodorou P., Grimm-Seyfarth A.and Onstein R.Full text 

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36. The effects of urbanisation on pollinators and pollination: A meta-analysis

Ecology Letters (2023), 00, 1– 14

Liang H., He Y., Theodorou P.* & Yang C. Full text 

*Corresponding author

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35. The degree of urbanisation reduces wild bee and butterfly diversity and alters the patterns of flower‑visitation in urban dry grasslands

Scientific Reports (2023), 13, 2702

Herrmann J., Buchholz S. & Theodorou P. Full text 

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​2022 (6)

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34. Bumble bee colony health and performance vary widely across the urban ecosystem

Journal of Animal Ecology (2022)

Theodorou P., Kühn O., Baltz L.M., Wild C., Rasti S.R., Bucksch C., Strohm E., Paxton R., Kurze C. Full text

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33. Anthropogenic effects on the body size of two Neotropical orchid bees

BMC Ecology and Evolution (2022)

Garlin J., Theodorou P., Kathe E., Quezada-Euan J., Paxton R.J., Soro A. Full text 

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32. How much does a drop of sugar solution benefit a hatchling of Chrysoperla pallida (Neuroptera Chrysopidae)?

Biological Control (2022), 104963

Pantaleoni R., Pusceddu M., Tauber C, Theodorou P., Loru L. Full text 

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31. The effects of urbanisation on ecological interactions

Current Opinion in Insect Science (2022), 100922

Thedorou P. Full text 

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30. Crop pesticide impacts reproductive male quality signals leading to reduction of mating in wild solitary bees

Journal of Applied Ecology (2022), 0 (1– 12)

Boff S., Conrad T., Raizer J., Wehrhahn M., Bayer M., Friedel A., Theodorou P., Schmitt T., Lupi D. Full text 

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​29. Global urban environmental change drives adaptation in white clover

Science (2022), 375:1275-1281

Santangelo J.S.,Ness R.W., Cohan B., Fitzpatrick C.R.,..............., Theodorou P.,......., Johnson M.T.J. Full text 

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2021 (7)

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28. An integrative environmental pollen diversity assessment and its importance for the Sustainable Development Goals

Plants, People, Planet (2021), 1-12

Hornick T., Bastl M., Bohlmann S.,......., Steffan-Dewenter I., Panagiotis P., Treudler R., Werchan B., Werchan M., Wolke R., Dunker S. Full text 

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​27. Honey bees increase social distancing when facing the ectoparasite Varroa destructor

Science Advances (2021), 7(44)

Michelina P., Alessandro C., Simona A., Emanuele S., Panagiotis T., Ignazio F., Alberto S. Full text

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26. A brief history and popularity of methods and tools used to estimate micro-evolutionary forces

Ecology and Evolution (2021), 00:1-21

Kidner J., Theodorou P., Engler J.O., Taubert M., Husemann M.. Full text

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25. Apple pollination is ensured by wild bees when honey bees are drawn away from orchards by a mass co-flowering crop, oilseed rape

Agricutlure Ecosystem and the Environment (2021), 315:107383

Osterman J., Theodorou P., Radzevičiūtė R., Schnitker P., Paxton R. Full text

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24. Urbanisation is associated with shifts in bumblebee body size with cascading effects on pollination

Evolutionary Applications (2021), 14: 53– 68

Theodorou P., Baltz L., Paxton R.J., Soro A. Full text

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23. Plant-pollinator networks in savannas of Burkina Faso, West Africa

Diversity (2021), 13(1)

Stein K., Coulibaly D., Balima L.H., Götze D., Linsenmair K.E., Porembski S., Stenchly K., Theodorou P. Full text

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22. A two-part modelling approach reveals a positive effect of pollinator biodiversity in boosting the pollination of apple flowers

Agricutlure Ecosystems and the Environment (2021), 306, 107197

Radzevičiūtė R., Theodorou P.*, Schlegel M., Paxton R. Full text

*corresponding author

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2020 (5)

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​​​21. Urban fragmentation leads to lower floral diversity, with knock-on impacts on bee biodiversity

Scientific Reports (2020), 10, 21756

Theodorou P., Herbst S., Kahnt B., Landaverde-González P., Baltz L., Osterman, Paxton R. Full text

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​​20. Disentangling the effects of local resources, landscape and climatic seasonality on bee biodiversity and plant-pollinator networks in tropical highlands

​Oecologia (2020), 194, 333-344

Escobedo-Kenefic N., Landaverde-González P., Theodorou P.*, Cardona E., Dardon M.J., Martínez O., Domínguez C.. Full text

*corresponding author

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19. Urban areas as hotspots for bees and pollination but not a panacea for all insects

Nature Communications (2020), 11(1), 1-13

Theodorou P., Radzevičiūte R., Lentendu G., Kahnt B., Husemann M., Bleidorn C., Settele J., Schweiger O., Grosse I., Wubet T., Murray T.E., Paxton R. Full text

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18. Population genetics of the European rabbit along a rural-to-urban gradient

Scientific Reports (2020), 10, 2448

Ziege M., Theodorou P., Jüngling H., Merker S., Plath M., Streit B., Lerp H. Full text

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17. Propolis consumption reduces Nosema ceranae infection of European honey bees (Apis mellifera)

Insects (2020), 11(2), 124

Mura A., Pusceddu M., Theodorou P., Angioni A., Floris I., Paxton R., Satta A. Full text

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2019 (4)

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16. Should I stay or should I go? Pollinator shifts rather than cospeciation dominate the evolutionary history of South African Rediviva bees and their Diascia host plants

Molecular Ecology (2019), 28:4118– 4133

Kahnt B., Hattingh W.N., Theodorou P., Wieseke N., Glennon K.L., van der Niet T., Paxton R., Goodman-Cron G.V. Full text

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15. The two prevalent genotypes of an emerging infectious disease,Deformed wing virus, cause equally low pupal mortality and equally high wing deformities in host honey bees

Viruses (2019), 11(2)

Tehel A., Vu Q., Bigot D., Gogol-Döring A., Koch P., Jenkins C., Doublet V., Theodorou P., Paxton R. Full text

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14. A roadmap for urban evolutionary ecology

Evolutionary Applications (2019), 12:384-398

Rivkin R., Santangelo J.S., Alberti M., ................., Diamond S.E.,......... Hendry A.,..... Losos J., MacIvor J.S.,.... McDonnell M., Miles L.S., Munshi-South J., Ness R.,....., Theodorou P., Thompson K.A.,.......Whitehead A., Winchell K.M., Johnson M.T.J. Full text

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13. Resin foraging dynamics in Varroa destructor infested hives. A case of medication of kin?  

Insect Science (2019), 26, 297-310

Pusceddu M., Piluzza G., Theodorou P., Buffa F., Ruiu L., Bullitta S., Floris I., Satta A. Full text

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​2018 (4)

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​​​12. Small and genetically highly structured populations in a long-legged bee, Rediviva longimanus, as inferred by pooled RAD-seq

BMC Evolutionary Biology (2018), 8:196

Kahnt B., Theodorou P., Soro A., Hollens H., Kuhlmann M., Pauw A., Paxton R. Full text

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11. The effects of raw propolis on Varroa-infested honey bee (Apis mellifera) workers

Parasitology Research (2018), 117, 3527-3535.

Pusceddu M., Floris I., Mura A., Theodorou P., Cirotto G., Piluzza G., Bullitta S., Alberto A., Satta A. Full text

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10. Genome-wide SNP scan suggests adaptation to urbanization in an important pollinator, the red-tailed bumblebee (Bombus lapidarius  L.)

Proceedings of the Royal Society B: Biological Sciences (2018), 285, 20172806.

Theodorou P., Radzevičiūte R., Kahnt B., Soro A., Grosse I., Paxton R. Author URL

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​9. Hopper parasitoids do not significantly benefit from non-crop habitats in rice production landscapes

Agriculture Ecosystems & Environment (2018), 254, 224-232.

Sann C., Theodorou P., Heong K.L., Villareal S., Settele J., Vidal S., Westphal W. Full text

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2017 (4)

 

8. Replication of honey bee RNA viruses across multiple bee species in apple orchards of Georgia, Germany and Kyrgyzstan

 

Journal of Invertebrate Pathology (2017),146, 12-23

Radzevičiūtė R., Theodorou P., Husemann M., Japoshvili G., Kirkitadze G., Zhusupbaeva A., Paxton R. Full text

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7. Sweat bees on hot chillies: provision of pollination services by native bees in traditional slash-and-burn agriculture in the Yucatan Peninsula of tropical Mexico

Journal of Applied Ecology (2017), 54, 1814-1824.

Landaverde-González P., Quezada-Euán J., Theodorou P., Murray T.E, Ayala R., Moo-Valle U., Husemann M., Vandame R., Paxton R. Full text.

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6. The structure of flower-visitor networks in relation to pollination across an agricultural to urban gradient

Functional Ecology (2017), 31, 838-847

Theodorou P., Albig K., Radzevičiūtė R., Settele J., Schweiger O., Murray T.E., Paxton R.. Full text.

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5. The population genetics of two orchid bees suggests high Ne, high dispersal, low diploid male production and only an effect of island isolation in lowering genetic diversity

Conservation Genetics (2017), 18, 607-619

Soro A., Quezada-Euan J., Theodorou P., Moritz R.F.A., Paxton R. Full text.

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2016 (1)

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4. Pollination services enhanced with urbanisation despite increasing pollinator parasitism

Proceedings of the Royal Society B: Biological Sciences (2016), 283, 20160561.

Theodorou P., Radzevičiūtė R., Settele J., Schweiger O., Murray T.E, Paxton R. Full text.

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2015 (1)

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3. A sting in the spit: widespread cross-infection of multiple RNA viruses across wild and managed bees

Journal of Animal Ecology (2015), 84, 615-624

McMahon D.P., Fuerst Matthias A., Jessica C., Theodorou P., Mark B.; Paxton R. Full text. 

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2013 (1)

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2. Habitat complexity affects how young of the year Atlantic cod Gadus morhua perceive predation threat from older conspecifics

Journal of Fish Biology (2013), 82, 2141-2146

Theodorou, P. , Snorrason, S. S. and Ólafsdóttir, G. Á. Full text.

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2012 (1)

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1. Reaching the limit: constrained behavioural flexibility of juvenile Atlantic cod (Gadus morhua) at current costal temperatures

Journal of Experimental Marine Biology & Ecology (2012), 413, 192-197

Theodorou, P. , Ólafsdóttir, G.Á. and Snorrason, S.S., Full text.

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Other

 

Linear mixed-effects models to control for the variability of microsatellite loci when comparing genetic diversity

Theodorou P. and Soro A. (2016). http://popgen.nescent.org/LMM-Genetic-Diversity.html (Vignettes for Population Genetics in R http://popgen.nescent.org)

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Baltz L. M., Osterman J., Benton F., Theodorou P., Mrozek J., Paxton R. (2020). Was summt und brummt in deutschen Apfelanlangen. Obstbau, 12, pp. 694-698.

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